Perona

Perona (ペローナ) is a member of the Mysterious Four lead by Warlord Gecko Moria and former Commander of the Wild Zombies and Surprise Zombies in Moria's Zombie army. She possesses the power of the Hollow-Hollow Fruit, a Paramecia Devil Fruit which allows her to produce ghosts known as Hollows, thus making her a Ghost Human. Her dream is to create a country where all the cute people are turned into zombie animals that obey only her. Her Negative Hollows allow her to depress people that come in contact with them making her a particularly dangerous foe, however during her battle with Usopp, she finds to her horror that her powers have no effect on him due to his nature pessimism, thus making him her natural enemy. She is defeated by Usopp and later tries to flee on the Straw Hats' ship, Thousand Sunny during the Special Zombie Oars rampage, only to encounter Bartholomew Kuma who uses his powers to send her to a island with a spooky castle after asking where she would like to go. She later befriends Roronoa Zoro when he is sent to the same island and the two discover that the castle is home to Zoro's rival, Dracule Mihawk. She stays on the island with Zoro and Mihawk during the Straw Hats two-years of training, nursing Zoro's injuries and later helps him reach Sabaody Archipelago to reunite with his crew and helped the Straw Hats in their escape by using her Negative Hollows on the Marines.

Role in Games
Perona first appears in One Piece: Pirate Warriors 2 as a playable character. During the games story, she is shown to be once again working with Gecko Moria but is defeated by the alliance between the Straw Hats, Jinbe, and the G-5 Marines under Smoker during their search for Luffy's crew. She later becomes separated from Moria after he decides to join Buggy's Great Pirate Alliance, Perona ends up working with Ex-Warlord Crocodile and the Arlong Pirates as part of the Arlong-Crocodile Alliance. However she dislikes working for Crocodile and after being defeated by the Straw Hat & Whitebeard Alliance forces, she decides to defect to their side and ends up joining them in their quest to restore Luffy crew that are still under the influence of the Frenzy Dial. She later returns with them to Thriller Bark, where she and the Straw Hat/Whitebeard Alliance finds Nico Robin and Usopp who are being controlled by Blackbeard and his ally Gecko Moria. To her delight she discovers that Usopp is no longer immune to her Negative Hollows as a side effect of being under the Frenzy Dial's influence and gleefully takes revenge on him by attack him with repeated Negative Hollow attacks. However her Negative Hollows ironically restores Usopp back to his true negative self restoring his immunity and freeing him from Blackbeard's control.

In One Piece: Pirate Warriors 3, Perona appears on Thriller Bark as Gecko Moria's subordinate. After she is defeated by the Straw Hats, she tries to flee during Oars attack but is defeated by the Straw Hats and is sent away to another island by Kuma.